For such an iconic work, the text of Origin was far from static. It was a living thing that Darwin continued to shape for the rest of his life, refining his ‘one long argument’ through a further five English editions. Many of his changes were made in response to comments in correspondence – with friends, strangers, supporters, and critics – and, taken together, letters and publications form a series of overlapping conversations with colleagues, reviewers, and the reading public.
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